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Publications

2024

Seath, JL., Firth, LB., Froneman, PW., Claassens, L. 2024. The use of gabions as an eco-engineering approach to enhance estuarine fish habitats in urbanised waterways. Ecological Engineering: 107431.

AK Whitfield & MKS Smith (2024) Future of the IUCN Endangered white steenbras Lithognathus lithognathus (Sparidae) – a tale of two estuaries, African Journal of Marine Science, 46:3, 155-167, DOI: 10.2989/1814232X.2024.2378714

Barnes, R.S.K., Seath, J.L. & Arendse, C.J. 2024. Differential sampling in the assessment of conservation and biodiversity merit: a comparison of the seagrass macrofauna in three nearby South African estuaries. Biodiversity and Conservation 33(8): 509-532. https://link.springer.com/article/10.1007/s10531-023-02754-0

Barnes, R.S.K. & Seath, J.L. 2024. The occurrence of a rich subtidal macrobenthic fauna in through-flow
residential marina canals, and the potential of such systems to be managed
as biodiversity assets. Ocean and Coastal Management 254: 107191 https://doi.org/10.1016/j.ocecoaman.2024.107191

2023

van Rensburg, H., Richoux, N.B. & Simon, C.A. 2023. The trophic position and isotopic niche of a cryptogenic tube-building polychaete in a protected clear-water estuarine bay. Estuarine, Coastal and Shelf Science 295: 108549. https://www.sciencedirect.com/science/article/pii/S0272771423003396?via%3Dihub

Schoeman, S. & Simon, C.A. 2023. Live to Die Another Day: Regeneration in Diopatra aciculata Knox and Cameron, 1971 (Annelida: Onuphidae) Collected as Bait in Knysna Estuary, South Africa. Biology 12(3): 483; https://doi.org/10.3390/biology12030483

Whitfield, A.K., Breen, C.M. & Read, M. 2023. Knysna Estuary—Jewel of the Garden Route.
Knysna Basin Project, Knysna. 381 pp.

2022

Barnes, R.S.K. 2022. Biodiversity differentials between seagrass and adjacent bare sediment change along an estuarine gradient. Estuarine Coastal and Shelf Science 274(S3):107951. https://www.sciencedirect.com/science/article/pii/S0272771422002098?via%3Dihub

Barnes, R.S.K., Claassens, L. & Seath, J. 2022. Where ecologically ’tis better to go brown than green: Enhanced seagrass macrobenthic biodiversity within the canals of a brownfield coastal marina. Biodiversity and Conservation 31(12): 2981-2997. https://link.springer.com/article/10.1007/s10531-022-02468-9

de Villiers, N.M., Harasti, D., Hodgson, A. & Claassens, L. 2022. The impact of a Reno mattress installation on an adjacent seagrass meadow and its macrofauna in a South African estuary. Regional Studies in Marine Science 52(1–2):102239. DOI: 10.1016/j.rsma.2022.102239

2021

Barnes, R.S.K. 2021. Patterns of seagrass macrobenthic biodiversity in the warm- temperate Knysna estuarine bay, Western Cape: a review. Aquatic Ecology. 55(6):1-19. https://link.springer.com/article/10.1007/s10452-021-09848-3

Claassens, L., de Villiers, N.M., Seath, J. & Wasserman, J. 2021. Distribution and density of the Critically Endangered estuarine pipefish across its range – implications for conservation. Aquatic Conservation: Marine and Freshwater Systems, 32(1): 28 – 41. https://onlinelibrary.wiley.com/doi/epdf/10.1002/aqc.3742 

Hodgson, A., Smith, F., Smith, P. & Claassens, L. 2021. Macrofauna associated with intertidal mussel beds in the Knysna estuarine embayment, South Africa. African Zoology 56(1):1-14. DOI:10.1080/15627020.2020.1848457

de Villiers, N.M., Harasti, D., Hodgson, A. & Claassens, L. 2021. A comparison of the fauna in eelgrass and erosion control structures in a warm temperate Southern African estuary. Regional Studies in Marine Science, 44(8):101757. DOI: 10.1016/j.rsma.2021.101757

Radloff, J., Hodgson, A. & Claassens, L. 2021. Settlement of the invasive mussel Mytilus galloprovincialis in a warm temperate estuarine embayment in South Africa. African Journal of Aquatic Science, 46(2):1-9. DOI: 10.2989/16085914.2020.1842170

2020

van Rensburg, H., Matthhee, C.A. & Simon, C. 2020. Moonshine worms ( Diopatra aciculata : Onuphidae, Annelida) in the Knysna Estuary, South Africa; taxonomy and distribution. Journal of the Marine Biological Association of the UK 100(6): 1-11. https://www.cambridge.org/core/journals/journal-of-the-marine-biological-association-of-the-united-kingdom/article/abs/moonshine-worms-diopatra-aciculata-onuphidae-annelida-in-the-knysna-estuary-south-africa-taxonomy-and-distribution/FE6D3C64D4264D29FF9681B3FAF78129

Barnes, R.S.K. & Claassens, L. 2020. Do beds of subtidal estuarine seagrass constitute a refuge for macrobenthic biodiversity threatened intertidally? Biodiversity and Conservation. 29(6): 3227-3244. https://link.springer.com/article/10.1007/s10531-020-02019-0

Wasserman, J., Claassens, L. & Adams, J. 2020. Mapping subtidal estuarine habitats with a remotely operated underwater vehicle (ROV). African Journal of Marine Science, 42(1): 123 – 128. https://www.tandfonline.com/doi/abs/10.2989/1814232X.2020.1731598

Claassens, L., Barnes, R.S.K., Wasserman, J., Lamberth, S., Miranda, N.F.A., van Niekerk, L. & Adams, J.B. 2020. Knysna Estuary health: ecological status, threats and options for the future. African Journal of Aquatic Science 45(1): 1-18. https://www.tandfonline.com/doi/abs/10.2989/16085914.2019.1672518

Claassens, L. & Harasti, D. 2020. Life history and population dynamics of an endangered seahorse (Hippocampus capensis) within an artificial habitat. Journal of Fish Biology, 97(4): https://doi.org/10.1111/jfb.14452

2019

Barnes, R.S.K. 2019. Abundance/occupancy/patchiness relations in estuarine seagrass macrobenthos. Estuarine, Coastal and Shelf Science. 228. https://www.sciencedirect.com/science/article/pii/S0272771419306870

De Villiers, N.M., Barker, C., Claassens, L., Hodgson, A.N. Conservation value of Codium tenue habitat for the Endangered Knysna seahorse Hippocampus capensis. Journal of Fish Biology. https://www.ncbi.nlm.nih.gov/pubmed/31621070

Barnes, R.S.K. 2019. Local patchiness of macrobenthic faunal abundance displays homogeneity across the disparate seagrass systems of an estuarine bay. Marine Environmental Research. 148: 99-107. https://doi.org/10.1016/j.marenvres.2019.05.001

Simon, C., du Toit, A.N., Smith, M.K.S., Claassens, L., Smith, F., Smith, P. 2019. Bait collecting by subsistence and recreational fishers in Knysna Estuary may impact management and conservation. African Zoology. 54(2): 91-103  https://doi.org/10.1080/15627020.2019.1608862

2018 

Barnes, R.S.K. 2018.  Context dependency in the effect of Ulva-induced loss of seagrass cover on estuarine macrobenthic abundance and biodiversity. Aquatic Conservation. In Press. https://doi.org/10.1002/aqc.2977

Hodgson, A.N., Claassens, L., Kandondi, S. 2018. Shell morphometrics and growth rate of the invasive bivalve mollusc Mytilus galloprovincialis in the Knysna estuarine embayment, South Africa. African Journal of Aquatic Science. In Press.

Claassens, L., Hodgson. 2018. Monthly population density and structure patterns of an endangered seahorse Hippocampus capensis: a comparison between natural and artificial habitats. Journal of Fish Biology. IN PRESS. https://www.ncbi.nlm.nih.gov/pubmed/29675915  

Claassens, L., Booth, A.J., Hodgson, A.N. 2018. An endangered seahorse selectively chooses an artificial structure. Environmental Biology of Fishes. 101(5): 723-733. https://link.springer.com/article/10.1007/s10641-018-0732-4  

Hamylton, S.M., Barnes, R.S.K. 2018. The effect of sampling effort on spatial autocorrelation in macrobenthic intertidal invertebrates. Hydrobiologia. 811:239–250. https://link.springer.com/article/10.1007/s10750-017-3491-x  

2017

Barnes, R.S.K., 2017. Little-known and phylogenetically obscure South African estuarine microgastropods (Mollusca: Truncatelloidea) as living animals. Journal of Natural History. 52: 87-113. http://www.tandfonline.com/doi/abs/10.1080/00222933.2017.1408867

Claassens, L., Hodgson, A.N., 2017. Gaining insights into in situ behaviour of an endangered seahorse using action cameras. Journal of Zoology. DOI: 10.1111/jzo.12509. http://onlinelibrary.wiley.com/doi/10.1111/jzo.12509/abstract

Pollard, M., Hodgson, A.N., Kok, H.M., Whitfield, A.K., 2017. Eelgrass beds and bare substrata – sparid and mugilid composition in contrasting littoral estuarine habitats. African Journal of Marine Science. 39(2): 211-224. http://www.tandfonline.com/doi/abs/10.2989/1814232X.2017.1333043

2016

Allanson, B.R., Human, L.R.D., Claassens, L., 2016. Observations on the distribution and abundance of a green tide along an intertidal shore, Knysna Estuary. Southern African Journal of Botany. 107, 49-53. http://dx.doi.org/10.1016/j.sajb.2016.02.197

Barnes, R.S.K., 2016. Spatial homogeneity of benthic macrofaunal biodiversity across small spatial scales. Marine Environmental Research. 122, 148-157. https://doi.org/10.1016/j.marenvres.2016.10.006

Claassens, L., 2016. An artificial water body provides habitat for an endangered estuarine seahorse species. Estuarine, Coastal and Shelf Science. 180. 1-10. http://dx.doi.org/10.1016/j.ecss.2016.06.011

Human, L.R.D., Adams, J.B., Allanson, B.R., 2016. Insights into the causes of an Ulva lactuca bloom and its impact in South Africa’s most important estuary. Southern African Journal of Botany. 107, 55-62. http://dx.doi.org/10.1016/j.sajb.2016.05.016

Pollard, M., Hodgson, A. N., 2016. Mytilus galloprovincialis (Mollusca: Bivalvia) in a warm-temperate South Africa estuarine embayment. African Journal of Aquatic Science. 41, 57-65. http://dx.doi.org/10.2989/16085914.2015.1119670

2015

Barnes, R.S.K., Hendy, I.W., 2015a.  Functional uniformity underlies the common spatial structure of macrofaunal assemblages in intertidal seagrass beds. Biological Journal of the Linnean Society. 115, 114-126.

Barnes, R. S. K., Hendy, I. W., 2015b.  Seagrass-associated macrobenthic functional diversity and functional structure along an estuarine gradient. Estuarine, Coastal & Shelf Science. 164, 233-243. http://dx.doi.org/10.1016/j.ecss.2015.07.050

Pinchuck, S.A., Allanson, B.R.and Hodgson, A.N., 2015. Evolutionary retention of defensive lateral pedal glands in the smallest siphonariid limpet (Gastropoda: Pulmonata). African Zoology. 50 (4), 327-330. http://dx.doi.org/10.1080/15627020.2015.1106925

2014

Allanson, B., Smith F., Smith P., 2014. The invertebrate macrobenthos of artificial rocky shores in the Knysna Estuary. Transactions of the Royal Society of South Africa. 69, 19-33.  http://dx.doi.org/10.1080/0035919X.2013.867910

Barnes, R.S.K., 2014. The nature and location of spatial change in species assemblages: a new approach illustrated by the seagrass macrofauna of the Knysna estuarine bay, South Africa. Transactions of the Royal Society of South Africa. 69, 75-80. http://dx.doi.org/10.1080/0035919X.2014.899277

Barnes, R. S. K., Barnes, M. K. S., 2014a. Biodiversity differentials between the numerically-dominant macrobenthos of seagrass and adjacent unvegetated sand in the absence of sandflat bioturbation. Marine Environmental Research. 99, 34-43. http://dx.doi.org/10.1016/j.marenvres.2014.05.013

Barnes, R. S. K., Barnes, M. K. S., 2014b. Spatial uniformity of biodiversity is inevitable if the available species are distributed independently of each other. Marine Ecology Progress Series. 516: 263-266.

Hodgson, A. N., Booth, A. J., David-Engelbrecht, V., Henninger, T. O., 2014. Some life-history parameters of the non-native amphipod Platorchestia platensis (Talitridae) in a warm temperate South African estuary. Transactions of the Royal Society of South Africa. 69, 97-106. http://dx.doi.org/10.1080/0035919X.2014.941961

2013

Barnes, R. S. K., 2013. Spatial stability of macrobenthic seagrass biodiversity. Marine Ecology Progress Series. 493, 127-139. https://doi.org/10.3354/meps10546

Barnes, R. S. K., 2013. Distribution patterns of macrobenthic biodiversity in the intertidal seagrass beds of an estuarine system, and their conservation significance. Biodiversity and Conservation. 22, 357 – 372.

Presentations

Lucienne R.D. Human, Adams J.B., Allanson B.R., The water quality in the Knysna Estuary: Status quo. Knysna Basin Project AGM 2017 https://knysnabasinproject.co.za/wp-content/uploads/2017/12/Dr-L-Human-The-water-quality-in-the-Knysna-Estuary.pdf

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About KERF

The Knysna Estuary Research Foundation is an NGO based in Knysna, South Africa, focused on coastal system research – particularly how these systems change and how communities can and should live within these systems. Our research is applied and aimed at guiding and facilitating management of our local systems, whilst developing novel conservation approaches for coastal systems in general. The Knysna Estuary Research Foundation is based on four pathways: Research, Educate, Conserve and Collaborate.

Recent Posts

MPA Day 2025: A Celebration of Connection, Community & Conservation in Knysna
Jessica SeathAugust 5, 2025
PRESS RELEASE 
Jessica SeathJuly 31, 2025

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Email: jessica@knysnaestuary.co.za

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